,*>LES ,D>W91 #A ,ISL&S1 & ,GALAPAGOS ,TORTOISES ,*>LES ,D>W9'S VOYAGE ( 5LIE F.D CLOSE/ TGR4 ,IF ,GOD _H CR1T$ EA* SPECIES AT ! 2G9N+ (! E>?1 :Y WD HE G 6ALL ! TR\# 6GR\P SIMIL> ORGANISMS ) EA* O!R8 ,? IS EXACTLY ! "Q A "Y1 UN"KN ,5GLI% G5TLEMAN F.D HMF ASK+ DUR+ ! #AHCJ;S4 ,L _M "Y P TD1 ,*>LES ,D>W9 0 UNSURE AB :AT HE WANT$ 6D #B ) 8 LIFE AF F9I%+ S*OOL4 ,HE F/ SET \ 6BE A DOCTOR1 B HE 0 S SQU1MI% AB BLOOD T HE RAN F ! CLASSROOM :5"E AN OP],N 0 2+ P]=M$4 ,9 8 %ORT YE> AT M$ICAL S*OOL1 ,D>W9 N"E!.S F.D ! "T 6JO9 A NATURAL HI/ORY SOCIETY1 ATT5D LECTURES ON OLD] EVOLU;N>Y !ORIES & 6LE>N TAXID]MY1 ! >T ( PRES]V+ BIOLOGICAL SPECIM5S = /UDY4 ,RESIGN$ 6MAK+ A LIV+ "SH[1 HE 5ROLL$ AT ! ,UNIV]S;Y ( ,CAMBRIDGE 6"W T[>DS A C>E] 9 LAW OR Z A P>SON1 ! L1D] (A SMALL ,5GLI% *UR*4 ,:ILE AT S*OOL1 ,D>W9 0 9TRODUC$ 6BEETLE COLLECT+ 0HIS C\S91 & LE>N$ A LOT AB NATURAL !OLOGY1 ! 2LIEF T BIOLOGICAL WOND] DEMON/RATES ! GLORY ( ,GOD4 ,( ALL ! EXOTIC LOC,NS /UDI$ 0BIOLOGI/S1 N"O >E Z FAM\S Z ! QUIRKY ,GALAPAGOS ,ISL&S1 ": ? "Y ,BRITI% MAN "ND ,*>LES #C ,D>W9 0 9SPIR$ 6"Q ": DI6]5T PLANT & ANIMAL SPECIES CAME F4 ,AL? HE DIDN'T R1LIZE X UNTIL HE RETURN$ 6,5GL& & %>$ 8 COLLEC;NS ) EXP]TS1 ! SIMPLE ECOSY/EMS & SLIW9'S NEW IDEA AB H[ EVOLU;N "W$1 IS A CLASSIC TALE (! WOND] ( 4COV]Y1 &A TE/A;T 6! P[] (A S+LE MAN'S M9D 9 FIGUR+ \ H[ ! NATURAL _W OP]ATES4 ,9 ,D>W9'S "D1 "! 7 NO PROFES.NAL SCI5TI/S4 ,9/1D1 SCI;E 0 MO/LY 3DUCT$ 0G5TLEM5 ) 9H]IT$ W1L? T DIDN'T NE$ A JOB4 ,P (! HI<] CLASSES EV5 W5T S F> Z 6LOOK D[N _! NOSES AT P T _H 6"W = _! LIV+4 ,2C HE ?"\ HE NE$$ ! M"OY1 #D ,D>W9 RELUCTANTLY />T$ LOOK+ =A JOB 9 ! *UR* AF 8 PLANN$ SCI5TIFIC VAC,N 6! ,CAN>Y ,ISL&S FELL "?4 ,9/1D1 HE RCVD A LR 9VIT+ HM 6S]VE Z -PANION & NATURALI/ ON A ,BRITI% NAVAL SURVEY (! COA/ ( ,S\? ,AM]ICA4 ,D>W9 JUMP$ AT 8 *.E1 8 UNCLE'S 5C\RAGE;T DEF1T+ ! PROTE/S ( 8 "F1 & SET \ ON ! F/ & ONLY GRT ADV5TURE ( 8 LIFE4 ,ASIDE F 8 DUTY 6KEEP ! CAPTA9 -PANY1 ,D>W9 TRAVEL$ UP & D[N ! COA/ ( ,S\? ,AM]ICA1 %OOT+ BIRDS & COLLECT+ BEETLES1 ID5TIFY+ ROCKS & UNE>?+ FOSSILS1 SPECULAT+ AB ! =M,N ( ISL&S & TAK+ PLANT SAMPLES4 ,D>W9 NOT$ T ! FOSSIL ANIMALS HE F.D 9 ,S\? ,AM]ICA M CLOSELY RESEM#D ^? LIV+ 9 ,S\? ,AM]ICA ?AN ANY": ELSE1 & HE 0 AL /RUCK )! SIMIL>;Y ( ISL& ANIMALS 6^? LIV+ CLOSE #E 0ON ! MA9L&4 ,^! ?+S PO9T$ 6A NATURAL PROCESS ( ANIMAL ORIG9S -- :Y WD ,GOD G 6ALL ! TR\# 6PLACE SIMIL> ANIMALS REM>KABLY CLOSE TGR8 ,! ,GALAPAGOS ,ISL&S ,LET'S TAKE A CLOS] LOOK AT ! ,GALAPAGOS &! UNIQUE ANIMALS T CALL ^! ISL&S HOME4 ,9 YR KIT Y W F9D A TEXTUR$ MAP &A BUN* ( PLA/IC ANIMAL MODELS4 ,IGNORE ! L>GE TORTOISES = N[1 B LET'S TAKE A LOOK AT ! MAP4 ,! ,GALAPAGOS >*IPELAGO SITS "R ON ! EQUATOR1 & 3SI/S ( SIX L>GE & Q A FEW SMALL] ISL&S4 ,TAKE A MO;T 6FAMILI>IZE YRF )! %APE & "NS (! ISL&S4 ,! ISL&S H ,SPANI% "NS1 2C !Y >E "P (! C.TRY ( ,ECUADOR1 ": ,SPANI% IS ! PRIM>Y LANGUAGE4 ,! ISL&S >E COV]$ 0VOLCANOES1 7,V4 ON ! MAP /&S = ,VOLCAN1 ! ,SPANI% ^W = VOLCANO7 & #F M.TA9S 7,SI]RA ON ! MAP74 ,! ISL&S 7 =M$ F A WE/W>D-MOV+ VOLCANIC HOT SPOT1 M1N+ T ! ISL&S >E MADE F COOL$ LAVA1 & T ! EA/]N ISL&S >E SIGNIFICANTLY OLD] ?AN ! WE/]N "OS4 ,! L>GE WE/]N ISL&1 ,ISLABELA1 IS S "Y T X / HAS M ?AN FIVE ACTIVE VOLCANOES T SPEW MOLT5 LAVA FIELDS ON A REGUL> BASIS4 ,TRY & GET A S5SE =! RELATIVE SIZE1 ORI5T,N1 POSI;N1 & 4T.E 2T ISL&S -- ? W 2COME IMPORTANT LAT]4 ,AL NOTICE T "S (! ISL&S H A MOI/ 9T]IOR SURR.D$ 0>ID S1COA/4 ,! MOI/ L&S >E AT HI<] ELEV,N1 & >E ECOLOGIC,Y V DI6]5T F ! DRY1 ROCKY L& NE> ! OC1N & ON ! SMALL] ISL&S4 ,! ,GALAPAGOS IS FAM\S = XS .5DEMIC .SPECIES1 ORGANISMS T O3UR NO": ELSE ON E>?4 ,P]H MO/ WELL-"KN >E ! M>9E IGUANAS -- ! ONLY SPECIES ( #G IGUANAS T REGUL>LY SWIM & FE$ "U ! WAT]4 ,! IGUANAS SLEEP1 BRE$1 & RELAX ON L&1 B V5TURE \ 96! SURF 6EAT S1WE$4 ,! IGUANAS >E .ECTO!RMS1 M1N+ T !Y GET _! H1T F ! 5VIRON;T4 ,2F DIV+ 9 ! COLD WAT] !Y M/ H1T UP 0BASK+ 9 ! SUNLI ! %OREL9E1 B ! L>G] MALES H 5 BODY MASS T !Y C SWIM F> (F%ORE & BACK 2F GETT+ TOO COLD ,9CLUD$ >E SMALL MODELS (! M>9E IGUANA & ANO!R 5DEMIC REPTILE -- ! ,GALAPAGOS L& IGUANA4 ,^! IGUANAS >E ! LIKELY ANCE/OR (! M>9E IGUANAS1 >E ADAPT$ 6EAT ! T\<1 LOCAL CACTUS1 & 7 ORIG9,Y S PL5TI;L T ,D>W9 -PLA9$ HE CDN'T PIT* 8 T5T )\T COV]+ "O ( _! BURR[S4 ,IF Y VISIT$ ! ,GALAPAGOS TD1 H["E1 Y'D #H H 6BE LUCKY 6SEE "O 9 ! WILD4 ,:Y D Y ?9K ? IS8 ,! ,GALAPAGOS IS AL HOME 6A CURI\S BIRD -- ! FLIGE & LUMB]+ L& TORTOISES T ,D>W9 BRIEFLY 5C.T]$ ON ,SANTA ,CRUZ ,ISL& 7CALL$ ,9DEFATIGA# 0! ,BRITI%7 _H BE5 "KN = DECADES 0:AL]S & PIRATES 6BE DELICI\S & 3V5I5T S\RCES ( M1T -- !Y CD 2 /OR$ 9 ! HULL (A SAIL+ %IP & WDN'T DIE = WEEKS4 ,D>W9 SOON R1LIZ$ T ? ABIL;Y CD AL ALL[ ! #I TORTOISES 6SURVIVE OC1N TRAVEL 2T ISL&S1 & EV5 EXPLA9 :Y ! TORTOISES 7 S SIMIL> 6! "OS HE OBS]V$ LIV+ 9 ,S\? ,AM]ICA4 ,9/1D ( 2+ PLAC$ ON EA* ISL& SPECIFIC,Y 0,GOD1 HE R1LIZ$1 !Y CD H FLOAT$ "! 0PREVAIL+ CURR5TS1 OR RAFT$ ON LOGS ( VEGET,N TORN F ! COA/ DUR+ /ORMS4 ,FEEL ! ,GALAPAGOS TORTOISE %ELL MODELS 9CLUD$ 9 ! KIT4 ,Y'LL NOTICE T "S ( !M H A SA4LE-%AP$ >RAIGN;T 7^! TORTOISES H AN >* 9 _! %ELL HI< OV] _! NECK1 %AP$ L A SA4LE =A HORSE71 "S H A HIE EXACTLY ! SAME4 ,"! >E ?IRTE5 SUBSPECIES OR RACES ( ,GALAPAGOS TORTOISE1 & EA* O3UPIES A DI6]5T GEOGRAPHICAL "P (! ISL&S4 ,"R N[1 SP5D A FEW M9UTES EXAM9+ EA* #AJ TORTOISE SUB-SPECIES MODEL 9 DETAIL4 ,MAKE A M5TAL LI/ (! SIMIL>ITIES & DI6];ES T Y NOTE1 & TRY 6IMAG9E H[ ^! DI6];ES CAME AB4 ,BE> 9 M9D T EA* TURTLE IS F.D ON ONLY "O1 OR J A FEW1 (! _M ,GALAPAGOS ,ISL&S4 ,:AT IS IMPORTANT AB ! PRES;E ( CLOSELY-RELAT$ B UNIQUE SPECIES LIV+ ON ISOLAT$ ISL&S LOCAT$ CLOSE TGR8 ,D>W9 F/ NOTIC$ ? PATT]N ( SLIE V SIMIL> 6^? ON ! MA9L&1 B H EVOLV$ A SLIG] BODY1 "W TGR 6HUNT & GA!R FOOD1 & >E M HESITANT 6FLY ?AN MA9L& SPECIES4 ,9 A4I;N1 ! MOCK+BIRDS ON ,ESPANOLA ,ISL& >E L>G]1 MANGI]1 & H A M CURV$ B1K4 ,SCI5TI/S /UDY+ ! S1BIRDS ON ? ISL& H NOTIC$ T ! #AA MOCK+BIRDS USE ? CURV$ BILL 6BR1K 96! E7S ( BOOBIES1 GULLS1 & ALBATROSS1 & EV5 6DR9K ! BLOOD ( S1BIRD *ICKS4 ,D>W9 IMMLY NOTIC$ ! SIMIL>ITIES & DI6];ES 9 ^! SPECIES & RECOGNIZ$ T ! GEOGRAPHIC PROXIM;Y M1NT "S?+ -- X DIDN'T MAKE S5SE T SPECIAL CRE,N WD PLACE ^! ANIMALS S CLOSE TGR4 ,X SEEM$ Z IF ! BIRDS _H MOV$ 6,ESPANOLA1 & !N DIV]G$ SLIW9 CAME BACK 6,5GL&1 HE %>$ 8 ,GALAPAGOS SPECIM5S ) .ORNI?OLOGI/S1 SCI5TI/S T SPECIALIZE 9 BIRDS1 :O QKLY NOT$ T ALL (! DULL BR[N_/BLACK BIRDS ,D>W9 COLLECT$ 7 N TOT,Y UNIQUE SPECIES Z ,D>W9 _H ORIG9,Y ?"\1 B 7 ACTU,Y ALL CLOSELY RELAT$ 6BIRDS CALL$ ,TANAG]S ON ! ,S\? ,AM]ICAN MA9L&4 ,! IMPLIC,NS1 Z #AB ,D>W9 R1LIZ$1 7 ENORM\S4 ,"O GR\P ( BIRDS _H COLONIZ$ A GR\P ( (F%ORE ISL&S & DEVELOP$ 96SEP>ATE TYPES ) UNIQUE BILLS ON DI6]5T ISL&S4 ,X DIDN'T MAKE S5SE = ,GOD 6PUT ANIMALS ON ! ISL&S 9 ? WAY -- :Y WDN'T HE J PUT ! SAME SPECIES ON ! ISL&S Z ON ! MA9L&8 ,:Y WD HE DELIB]ATELY >RANGE ! PLACE;T (! BIRDS 6MAKE X SEEM L !Y 7 RELAT$ 6EA* O!R8 ,NO?+ MADE ANY S5SE AT ALL 6,D>W9 UN.S A SMALL GR\P ( BIRDS _H TRAVEL$ 6! ,GALAPAGOS & !N RADIAT$ 96SEP>ATE TYPES ALL ON _! [N4 ,BACK ,HOME & ,EUREKA6 -- ,NATURAL ,SELEC;N ,D>W9 R1LIZ$ T ! BIRDS EVOLV$ F A S+LE GR\P ALM AT ONCE1 B HE CDN'T FIGURE \ H[ X HAPP5$4 ,H[ CD "O TYPE ( ANIMAL ADAPT 6A NEW HABITAT & 2COME ANO!R TYPE8 ,! #AC 9SPIR,N = 8 ANSW] CAME F A SEEM+LY UNRELAT$ ESSAY ON POPUL,NS 0! MA!MATICIAN ,?OMAS ,MAL?US4 ,MAL?US R1SON$ T IF ! HUMAN POPUL,N KEPT 9CR1S+ AT XS CURR5T RATE1 "! WD EV5TU,Y 2 M P ?AN WE CD GR[ FOOD 6FE$1 & MO/ BABIES WD N SURVIVE 62COME ADULTS4 ,D>W9 SUPPOS$ ? 0 AL ! CASE 9 WILD ANIMALS1 & ?US HIT ^U ! IDEA ( NATURAL SELEC;N4 ,PUT SIMPLY1 X SU7E/S T 2C ONLY A C]TA9 NUMB] ( ANIMALS F EA* G5],N C SURVIVE1 & 2C "! >E SMALL DI6];ES )9 POPUL,NS 7"S ROB9S H "E S SLIG] B1KS1 = EXAMPLE7 ONLY ! "OS MO/ ADAPT$ 6_! 5VIRON;T WD 2 A# 6GR[ 6ADULTHOOD & H *N ( _! [N4 ,? ME*ANISM CD F9,Y EXPLA9 ! *ANGE ( SPECIES OV] "T -- ! ANIMALS BE/ ADAPT$ 6_! HABITAT WD PASS ON _! "*I/ICS 6FUTURE MEMB]S (! SPECIES4 #AD ,AL"R1 LET'S TRY AN ACTIV;Y 6GET A BETT] FEEL = H[ NATURAL SELEC;N RE,Y "WS4 ,9 YR KIT Y'LL F9D A TRAY & "S WOOD5 B1DS4 ,"S ( !M >E FUZZY1 & "S SMOO?4 ,9 ? GAME1 ALL (! B1DS >E ANIMALS 9 ! SAME SPECIES CALL$ ,SQUASSUMS1 &! DI6];E 9 TEXTURE REPRES5TS DI6]5T TRAITS PRES5T 9 ! POPUL,N4 ,IMAG9E T \R POPUL,N ( ,SQUASSUMS 0 LIV+ 9 A PRETTY OP5 5VIRON;T1 BRE$+ .ASEXU,Y1 M1N+ T "O SQUASSUM J DIVIDES 96TWO NEW SQUASSUMS1 & 7 2+ T]RORIZ$ ) REGUL> PRED,N 0,CLAW/]S 7Y GET 6PLAY ! CLAW/]74 ,=TUNATELY =! ,SQUASSUMS1 A FUZZY GR.D PLANT CALL$ ,GRABWE$ HAS BE5 GR[+ 9 _! HABITAT 7X'S :AT MAKES ! TRAY FUZZY71 PROVID+ PROTECTIVE COV] = "S ( \R ,SQUASSUMS4 ,:5 ! #AE GRABWE$ />T$ GR[+1 HALF (! ,SQUASSUMS 7 FUZZY1 & HALF 7 SMOO?4 ,LET'S SEE :AT HAPP5S AF A FEW G5],NS4 ,/>T \ 0PLAC+ T5 FUZZY & T5 SMOO? ,SQUASSUMS 9 ! HABITAT4 ,E G5],N Y'LL SIMULATE A ,CLAW/] ATTACK 0HOLD+ YR H& 9 ! %APE (A GIANT CLAW & DRA7+ X ACR ! 5VIRON;T 6GRAB #B-C SQUASSUMS4 ,AF T1 Y'LL SIMULATE ! GR[? (! ,SQUSSUM POPUL,N 0GRA2+ #E TOT,Y R&OM SQUASSUMS F ! HABITAT & REPLAC+ EA* "O Y GRAB )! ORIG9AL & TWO O!R'S J L X -- J Z IF EA* FUZZY SQUASSUM _H TWO FUZZY *N1 & EA* SMOO? SQUASSUM _H TWO SMOO? *N4 ,RUN ! GAME = #AJ G5],NS 7EA* DIVID+ POPUL,N & CLAW/] ATTACK IS "O G5],N7 & !N C.T ! SQUASSUMS LEFT 9 ! HABITAT4 ,H[ _M SMOO? & FUZZY SQUASSUMS >E "!8 ,>E ! NUMB]S #AF DI6]5T F ! 2G9N+ 7"! 7 #AJ ( EA* 9 ! 2G9N+78 ,IF !Y >E DI6]5T1 :AT D Y ?9K CAUS$ ! DI6];ES8 ,LET'S GET BACK 6\R L& TORTOISES4 ,IF NATURAL SELEC;N CAUS$ ! TORTOISES 6DEVELOP UNIQUE %ELL %APES ON EA* ISL&1 "! M/ H BE5 A WAY 9 : ^! DI6];ES CD B5EFIT EA* SUB-GR\P ( TORTOISE1 J L H[ ! FUZZY COAT ON "S (! ,SQUASSUMS ALL[$ !M 6AD"H )! ,GRABWE$ & AVOID PRED,N F ! ,CLAW/]S4 ,BIOLOGI/S 2LIEVE T ! SA4LE- %AP$ %ELL DESIGN HELPS TORTOISES 9 ! DRI] ISL&S 6R1* VEGET,N \ ( R1* 6TORTOISES )A NORMAL DOM$ %ELL4 7,FEEL H[ ! SA4LE FL>E ABV ! NECK REMOVES ! "P (! %ELL T WD O!RWISE BLOCK ! NECK F 2COM+ TOT,Y V]TICAL74 ,! POPUL,NS ON ^! ISL&S MAY H />T$ \ ) #AG NORMAL DOME %APES1 B EVOLV$ SA4LE-BACK$ %ELLS DUR+ "TS ( DR"\1 A P]FECT EXAMPLE ( ,D>W9'S NATURAL SELEC;N !ORY4 ,NOTICE T ! .HOOD5SIS TORTOISE1 "O (! MO/ SA4LEBACK$ ( ALL1 -ES F ,ESPANOLA1 "O (! DRY ISL&S1 & T .PORT]I1 "O (! MO/ DOM$ TORTOISES1 -ES F MOI/ ,SANTA ,CRUZ ,ISL&4 ,:5 FOOD 0 SC>CE1 ! TORTOISES ) %ELLS T ALL[$ !M 6R1* HI<] GOT M 6EAT1 SURVIV$ AT A HI<] RATE ?AN ! DOM$ TORTOISES1 & GAVE RISE 6M SA4LE-%AP$ (FSPR+4 ,? PROCESS1 ": "O TYPE ( ANIMAL DIV]SIFIES 96_M DI6]5T TYPES T SPECIALIZE 9 DI6]5T ECOSY/EMS IS CALL$ .ADAPTIVE .RADI,N4 ,AL"R1 WE H A HYPO!SIS = :Y "S TORTOISES >E SA4LEBACKS & "S >E N -- ! SA4LEBACK =M EVOLV$ ON DRI] ISL&S ": FOOD IS LESS AVAILA# &! TORTOISES NE$ 6BE A# 6R1* HI<] = #AH ANY AVAILA# VEGET,N4 ,B G BACK 6YR NOTES F 2F -- >E ^! ! ONLY %APE *ANGES Y NOTIC$8 ,IF Y 7 HIATE SPECIES1 ! ORIG9AL SPECIES IS F/ USU,Y SEP>AT$ 96TWO OR M POPUL,NS 0"S SORT ( B>RI]4 ,! B>RI] IS USU,Y A #AI PHYSICAL /RUCTURE1 S* Z A M.TA9 RANGE OR A DES]T4 ,AF A:ILE1 R&OM *ANGES A3UMULATE 9 EA* POPUL,N MAKE !M TOO DI6]5T F EA* O!R 6REPRODUCE & PRODUCE VIA# (FSPR+1 RESULT+ 9 .SPECI,N1 OR ! =M,N ( NEW SPECIES4 ,? A3UMUL,N ( R&OM *ANGES 2T TWO SPECIES IS "KN Z .G5ETIC .DRIFT1 & PROBABLY EXPLA9S :Y ! ,GALAPAGOS TORTOISE SUBSPECIES1 AL? !Y >E \TW>DLY V SIMIL>1 _C BRE$ TGR TD4 ,! E6ECT IS SIMIL> 6H[ ! SEP>,N (! ,ATLANTIC ,OC1N HAS CAUS$ ! ,5GLI% LANGUAGE 6BE SPOK5 )A DI6]5T A35T 9 ! ,UNIT$ ,/ATES &! ,UNIT$ ,K+DOM4 ,! ,AM]ICAN A35T DEVELOP$ 0! A3UMUL,N ( R&OM *ANGES1 DOESN'T MAKE X EASI] 6-MUNICATE 9 ! ,UNIT$ ,/ATES ?AN 9 ,5GL&1 & IF ! ,,USA & ,5GL& 2CAME TOT,Y SEP>AT$1 CD EV5TU,Y #BJ 2COME S 4T9CT T ,5GLI%-SP1K]S F ! TWO C.TRIES CD N "U/& EA* O!R4 ,LET'S PLAY ANO!R GAME 6BETT] "U/& H[ ? CD H HAPP5$4 ,TAKE \ ! POK] *IP MON/]S -- NOTE T "! >E DI6]5T TYPES -- #AJ EA* ( MON/]S ,A1 ;,B1 & ,C4 ,?R[ ALL ( !M ON ! TA# & SCATT] !M >.D4 ,NEXT1 WE'LL PRET5D A M.TA9 RANGE GREW UP 9 ! MI4LE (! TA#4 ,)\T FEEL+ 6SEE : TYPE ( MON/]S Y'RE GRA2+1 SEP>ATE ! MON/]S 96?REE GR\PS4 ,NEXT1 C.T ! TYPES ( MON/]S LEFT 9 EA* SMALL] GR\P4 ,DOES EA* GR\P ALL H ! SAME NUMB] ( EA* TYPE ( MON/]8 ,:Y N8 ,:AT DOES ? M1N =! FUTURE (! POK] *IP MON/]S8 ,IF Y 7 A ,GALAPAGOS TORTOISE ON "O (! ISL&S 9 ! ,GALAPAGOS1 "! WD 2 TWO MA9 B>RI]S 6YR MAT+ ) ANY #BA O!R ,GALAPAGOS TORTOISE -- ! OC1N1 & IMPASSA# FIELDS ( RAZOR-%>P LAVA4 ,BIOLOGI/S 2LIEVE T ! ANCE/RAL TORTOISE POPUL,N L&$ ON EI ,ESPANOLA OR ,SAN ,CRI/OBAL ISL& 7! ISL&S CLOSE/ 6! MA9L&71 & !N =TUIT\S CURR5TS OR1 _M ?\S&S ( YE>S LAT]1 SAILORS TRANSPORT$ TORTOISES F ISL& 6ISL&4 ,^! EV5TS 7 EXTREMELY R>E1 B GIV5 ! L;G "T SCALES 9VOLV$ 7TORTOISES H BE5 ON ! ISL&S = AT L1/ #B MILLION YE>S7 >E ALM C]TA9 6H O3URR$4 ,ONCE A POPUL,N GOT />T$ \ ON A NEW ISL&1 X 0 TOT,Y ISOLAT$ = ALL PRACTICAL PURPOSES4 ,"! 7 N 5 TORTOISES A3ID5T,Y FLOAT+ BACK & =? 6KEEP ! TWO GR\PS 3NECT$1 &! POPUL,NS DIV]G$4 ,ANO!R CAUSE 9 : TORTOISE POPUL,NS CD 2 SEP>AT$ IS ! RELATIVELY ACTIVE GEOLOGY (! .>*IPELAGO1 OR ISL& #BB GR\P4 ,EXAM9E ! L>GE/ ISL& ON YR MAP1 ,ISABELLA1 : AL HAPP5S 6BE "O (! "YE/4 ,! ISL& HAS FIVE MAJOR VOLCANOES1 ,VOLCANS ,WOLF1 ,D>W91 ,ALC$O1 ,SI]RA ,NEGRA1 & ,AZUL4 ,:5 ^! VOLCANOES ERUPT1 MOLT5 HOT LAVA FL[S \ (! CRAT]S1 COV]+ ! ISL&S 9 ?ICK SPR1DS ( %>P ROCK1 /]ILIZ+ ! 5VIRON;T & CR1T+ AN IMPASSI# Z"O = TORTOISES4 ,N ONLY IS ! REGION A NAVIG,NAL *ALL5GE 6! SL[- MOV+ TORTOISES1 B ! NEW LAVA FL[S >E DEVOID ( ALL PLANT LIFE = _M YE>S1 & "!=E "!'S NO FOOD =! TORTOISES 9 ^! REGIONS4 ,"S"TS1 2C ! HABITAT ON EI SIDE (! B>RI] IS DI6]5T1 POPUL,NS "UGO SPECI,N Z ! RESULT ( ADAPTIVE RADI,N & G5ETIC DRIFT "W+ TGR4 ,TAKE ANO!R LOOK AT ! MAP (! ,GALAPAGOS ,ISL&S -- C Y #BC IMAG9E H[ ? SORT ( EVOLU;N CD O3UR "H8 ,EVOLU;N>Y ,HI/ORY ( ,GALAPAGOS ,TORTOISES ,WE'VE N[ 4CUSS$ TWO ME*ANISMS T C EXPLA9 ! DI6];E 9 TORTOISE %ELLS 9 ! ,GALAPAGOS -- ADAPT,N 6! LOCAL 5VIRON;T1 & SPECI,N F ISOL,N1 CALL$ ALLOPATRIC SPECI,N4 ,FEEL ! MODELS Y'VE BE5 GIV51 & KEEP+ ^! TWO FACTORS 9 M9D1 TRY 6FIGURE \ AN EVOLU;N>Y HI/ORY (! TORTOISE SPECIES4 ,GRAB ! MAP AG -- Y'LL NE$ 6"K ": ! TORTOISES >E F.D ON ! ISL&S3 ,BECKI ,VOLCAN ,WOLF1 ,ISABELA ,*ATHAM5SIS ,SAN ,CRI/OBAL ,GUN!RI ,SI]RA ,NEGRA1 #BD ,ISABELA ,HOOD5SIS ,ESPANOLA ,PORT]I ,SANTA ,CRUZ ,>E Y HAV+ TR\# FIGUR+ X \8 ,X'S PRETTY DI6ICULT4 ,BIOLOGI/S H BE5 /RU7L+ 6FIGURE \ ! L91GE ( ^! TORTOISES = _M YE>S1 & ONLY REC5TLY HAS ,,DNA EVID;E BE5 US$ 6%$ LI*IPELAGO _H 6BE DOME-%AP$1 2C ALL (! TORTOISES ON ! MA9L& ( ,S\? ,AM]ICA >E DOME-%AP$4 ,! F/ TORTOISES PROBABLY L&$ ON ! ISL& ( ,ESPANOLA OR ,SAN ,CRI/OBAL1 ! CLOSE/ ISL&S 6! MA9L&1 & AL ! OLDE/ 7REMEMB] -- WE R1D AB ! MOV+ HOT SPOT &! =M,N (! ,GALAPAGOS ,ISL&S74 ,! TORTOISES ON ,SAN ,CRI/OBAL H A %ELL %APE 9T]M$IATE #BE 9 2T ! DOM$ & SA4LE %AP$ V>IETIES1 & ^? ON ,ESPANOLA >E AM;G ! MO/ EXTREME SA4LE %AP$ RACES4 ,F "H1 TORTOISES MOV$ 6! RE/ (! ISL&S 0PREVAIL+ W9DS & CURR5TS4 ,"! IS EVID;E T A N[-EXT9CT SUB-SPECIES ( DOM$ TORTOISES 0 E/ABLI%$ ON ,SAN ,CRI/OBAL1 M1N+ T X IS ! SITE ( ORIG9AL COLONIZ,N4 ,? 4CUS.N BR+S UP AN IMPORTANT PO9T AB ISL&S -- _! =M,N4 ,ISL&S C =M 9 TWO WAYS3 EI 0SEP>AT+ F A 3T95T & FLOAT+ \ 6SEA1 Z 9 ! CASE ( ,MADAGASC>1 OR 2+ =M$ Z "P (A UNIQUE VOLCANIC EV5T -- _M ISL&S >E ACTU,Y J ! TIPS ( HUGE "UWAT] VOLCANOES4 ,2LIEVE X OR N1 ! VOLCANO T =MS ! ISL& ( ,HAWAII IS ACTU,Y ! TALLE/ M.TA9 ON E>?1 /RET*+ OV] #AJ1JJJ FEET HI<] F ! FLOOR (! OC1N ?AN ,"EE/ RISES ABV ! L&4 ,:5 VOLCANIC ISL&S S* Z #BF ! ,GALAPAGOS & ,HAWAII =M1 !Y'RE 9ITI,Y DEVOID ( LIFE4 ,"EY?+ LIV+ ON ^! ISL&S N[ 7EXCEPT = SPECIES T HUMANS H A3ID5T,Y OR PURPOSE;LLY 9TRODUC$7 _H 6MAKE X 6! ISL&S 0*.E4 ,9 ! #AIGJ;S1 ! VOLCANO ,KRAKATAUA EXPLOD$ 9 ! ,9DIAN ,OC1N 9 "O (! L>GE/ VOLCANIC ERUP;NS "E RECORD$4 ,Z "P (! ERUP;N1 A NEW ISL& ( FRE% LAVA 0 =M$1 CALL$ ,ANAKRAKATAU1 OR SON ( ,KRAKATAU4 ,N[-FAM\S BIOLOGI/S1 9CLUD+ ,E4,O4 ,WILSON1 JUMP$ AT ! *.E 6WAT* A NEW 5VIRON;T 2+ =M$ 2F _! EYES1 & DOCU;T$ ! SPECIES L&+ ON ! ISL& OV] "T4 ,! GR[? />T$ \ SL[1 A FEW SPECIES A MON?1 B !N 2GAN 6A3EL]ATE4 ,EV5TU,Y1 AN 5VIRON;T REM9ISC5T (! OLD] ,KRAATAUA DEVELOP$ & AF A :ILE1 ! TOTAL NUMB] ( SPECIES />T+ REMA9+ ! SAME1 EV5 ?\< NEW SPECIES KEPT %[+ UP4 #BG ,H[ CD ? BE8 ,WELL1 X TURNS \ "S (! SPECIES 7 GO+ EXT9CT4 ,! 4T.E_/>EA E6ECT #BH ,AF AN ISL& -ES 6EQUILIBRIUM1 ! NUMB] ( SPECIES LIV+ ON ! ISL& /AYS RELATIVELY ! SAME1 B X'S N NECESS>ILY ! SAME ASSEMBLY ( SPECIES ="E4 ,9 A4I;N1 SCI5TI/S H NOTIC$ T DI6]5T ISL&S H DI6]5T NUMB]S ( SPECIES4 ,6"U/& :Y1 X'S IMPORTANT AG 6?9K AB GEOGRAPHY4 ,TWO MA9 FACTORS A6ECT ! NUMB] ( SPECIES ON A GIV5 ISL&3 ! IMMIGR,N RATE 7H[ (T5 NEW SPECIES 2COME E/ABLI%$ ON AN ISL&71 &! EXT9C;N RATE 7H[ (T5 E/ABLI%$ SPECIES DIE (F74 ,:AT D Y ?9K 9FLU;ES ^! FACTORS8 ,F/1 ! IMMIGR,N RATE DEP5DS ON ! 4T.E 2T ! ISL& & O!R L& MASSES4 ,? MAKES S5SE1 2C ! FUR!R AN ISL& IS F A 3T95T1 ! LESS LIKELY X IS T ANIMALS & PLANTS W 2 LUCKY 5 6L& "!4 ,IF Y FIRE A %OTGUN AT A #BI T>GET #AJ FEET AWAY1 A LOT M (! PELLETS >E GO+ 6HIT ?AN IF YR T>GET IS #EJ FEET AWAY1 & ? EXPLA9S :Y ,MADAGASC> GETS A LOT M NEW SPECIES ?AN ANY": 9 ! ,HAWAIIAN ,ISL&S4 ,! BI7E/ FACTOR A6ECT+ ! EXT9C;N RATE IS SIMPLY ISL& SIZE -- BI7] ISL&S C SUPPORT M SPECIES1 2C "! >E M NI*ES ON L>G] ISL&S &! *.E ( EA* ORGANISM HITT+ R\< "TS & DY+ (F IS LESS5$ ,X'S SORT ( SIMIL> 6H[ L>G] CITIES H M DIV]SE RE/AURANTS -- SMALL T[NS DON'T H 5 D9]S 6SUPPORT _M DI6]5T K9DS ( FOOD OP;NS4 ,PULL \ ! *>T (! SPECIES- >EA-4T.E CURVE4 ,! RATE ( EXT9C;N IS PLOTT$ AG/ ! "R AXIS1 & RATE ( IMMIGR,N PLOTT$ AG/ ! LEFT AXIS4 ,?9K AB :AT ! *>T IS SAY+ AB ! TOTAL NUMB] ( SPECIES LIKELY 6BE PRES5T ON ANY GIV5 ISL&4 ,:AT K9D ( ISL& WD SUPPORT ! GRTE/ NUMB] #CJ ( SPECIES8 ,3SID] AN ISL&4 ,Y'RE PROBABLY PICTUR+ A SMALL PIECE ( L& 9 ! MI4LE (! OC1N ) G5TLE WAVES & PALM TREES4 ,9/1D1 IMAG9E C5TRAL P>K 9 ,,NYC4 ,X'S SORT ( L AN ISL&1 ISN'T X8 ,X'S A SMALL PIECE ( NATURAL >EA SURR.D$ 0! OC1N ( URBANIZ,N T IS ,NEW ,YORK ,C;Y4 ,IF AN ANIMAL WANTS 6GET 9 OR \ ( C5TRAL P>K1 X HAS 6CROSS BLOCKS ( D[NT[N C;Y TRA6IC -- Q A F1T =A RA3OON OR A MOOSE4 ,BIRDS & 9SECTS C R1DILY FLY 96! P>K F O!R NATURAL >1S CLOSE 6! C;Y1 B :AT IF ,NEW ,YORK ,C;Y 0 M* L>G]8 ,UNIQUE;S ( ISL& FAUNA ,ISL&S H REMA9$ IMPORTANT 9 BIOLOGY EV5 AF ! "DS ( ,D>W9 & ,WALLACE4 ,9DE$1 A :OLE SUBSET ( BIOLOGY1 CALL$ .ISL& #CA .BIOGEOGRAPHY1 HAS BE5 />T$ 6"U/& ! 4TRIBU;N ( PLANTS & ANIMALS ON ISL& SY/EMS4 ,F/ (F1 S"EAL CURI\S E6ECTS ( ISL&S ON ANIMAL SPECIES H BE5 NOT$4 ,6/>T1 ISL& SPECIES SEEM 62COME EI M* L>G] OR M* SMALL] ?AN _! MA9L& ANCE/ORS4 ,ON ! ,GALAPAGOS1 ! TORTOISES >E GIGANTIC1 WEI<+ UP 6#HHJ P.DS & GR[+ UP 6#F FEET 9 L5G?1 M* BI7] ?AN _! MA9L& ANCE/ORS4 ,C Y ?9K ( AN EXPLAN,N =! GIGANTISM ON ISL&S8 ,"! >E _M O!R EXAMPLES1 9CLUD+ ! GIANT ,GALAPAGOS C5TIP$ES1 &! L>G] MICE & RATS F.D ON ! ,*ANNEL ,ISL&S 9 ,CALI=NIA4 ,Z SCI;E WRIT] ,DAVID ,QUAMM5 HAS NOT$1 8,ISL&S >E HAV5S & BRE$+ GR.DS =! UNIQUE & ANOMAL\S4 ,!Y >E NATURAL LABORATORIES ( EXTRAVAGANT EVOLU;N>Y EXP]I;T,N40 ,ANIMALS ON ISL&S T5D 6EVOLVE V #CB BIZ>RE T5D5CIES4 ,! FAM$ M>9E IGUANAS (! ,GALAPAGOS >E A FANTA/IC EXAMPLE4 ,N ONLY D !Y GR[ 6SIZES F> 9 EXCESS ( _! MA9L& ANCE/ORS1 !Y H ADAPT$ 6SP5D A LOT ( "T 9 ! OC1N1 & >E ! ONLY "KN AQUATIC IGUANAS 9 ! _W4 ,:Y D Y ?9K ISL&S >E GD PLACES = WEIRD ANIMALS 6EVOLVE8 ,FUR!RMORE1 ISL& SPECIES (T5 LOSE _! CAPAC;Y = 4P]SAL4 ,MO/ ANIMALS & PLANTS H A LOT ( ABILITIES 9 ORD] 6SPR1D \ OV] ! L&SCAPE3 PLANTS H SE$S T GET /UCK ONTO ! HAIR ( ANIMALS & GET C>RI$ 6NEW LOC,NS1 & BIRDS C EASILY FLY 6NEW PLACES4 ,ON ISL&S1 H["E1 ? SEEMS 6BE REDUC$4 ,_M ISL& BIRDS 2COME FLIE USU,Y DEVOID ( SPECIES 9 ! 2G9N+1 ESPECI,Y VOLCANIC ISL&S L ! ,GALAPAGOS1 "! >E USU,Y ALM NO -PETITOR SPECIES PRES5T2 ANIMALS T "SH[ MAKE X 6ISL&S >E REL1S$ F ALL (! PRESSURES !Y EXP]I;ED AT HOME4 ,BIRDS H FEW] DANG]S 6FLY AWAY F1 & AT ! SAME "T "! >5'T A LOT ( PLACES 6FLY TO -- ! ANIMALS >E ISOLAT$ ON A SMALL PAT* ( L&4 ,"!=E1 )! MAJOR B5EFITS ( FLIE NO NATIVE WOODPECK]S 9 ! ,GALAPAGOS1 A SPECIES ( ,GALAPAGOS F9* HAS EVOLV$ F 2+ A CLASSIC SE$-EAT] 6PROB+ CACTUS PLANTS ) XS BILL & CACTUS SP9ES 6SEEK \ 9SECTS4 ,! F9* ISN'T Z E6ECTIVE 9 ? JOB Z A CLASSIC WOODPECK]1 B X GETS AL;G 9 ! ABS;E ( -PETITORS4 ,SIMIL>LY1 9 ! ABS;E ( GOATS & O!R MAMMAL H]BIVORES1 ! TORTOISES 9 ! ,GALAPAGOS 7 A# 6EVOLVE GIGANTIC SIZE 62COME ! DOM9ANT PLANT-EAT]S ON ! ISL&S4 ,Z A RESULT1 WE C SEE T ISL&S C ESS5TI,Y FUNC;N Z BIODIV]S;Y FACTORIES4 .,BIODIV]S;Y IS A M1SURE (! V>I,N ( LIFE =MS 9 AN ECOSY/EM OR GEOGRAPHIC REGION1 & C 2 BAS$ (F ! TOTAL V>I,N 9 #CE SPECIES1 9DIVIDUALS1 -MUNITIES1 OR G5ES 9 ! >EA4 ,2C EA* ISL& IS SEP>AT$ F E O!R1 EA* "O C EVOLVE XS [N UNIQUE SPECIES F.D NO": ON E>?1 3TRIBUT+ 6BIODIV]S;Y4 ,REMEMB] H[ ! ,GALAPAGOS TORTOISES 7 A# 6DIV]GE 96SUB- SPECIES J 2T ! SEP>ATE ISL&S 9 ! >*IPELAGO8 ,WELL1 ? SAME PROCESS O3URS _WWIDE1 ) E UNIQUE ISL& ECOSY/EM G5]AT+ TONS ( .5DEMIC .SPECIES 7^? SPECIES F.D NO": ELSE74 ,^! SPECIES >E (T5 SPECIALI/S1 WELL-ADAPT$ 6A V SPECIFIC WAY ( LIFE1 ) ADAPT,NS T ALL[ !M 6EXTRACT A WAY ( LIFE F A V SPECIFIC BIOLOGICAL NI*E4 ,A GRT EXAMPLE ( ? IS ! MANGROVE F9*1 "O ( ,D>W9'S F9*ES1 : HAS ADAPT$ 6PLUCK 9SECTS F ! ISOLAT$ COA/AL MANGROVE =E/S ON ! ,GALAPAGOS ,ISL&S ( ,F]N&9A & ,ISABELA4 #CF ,UN=TUNATELY1 ANIMALS T 2COME SPECIALIZ$ 6EXPLOIT A V N>R[ HABITAT (T5 2COME -PLETELY DEP5D5T ON A R>E ECOSY/EM & >E "!=E HILY W5T EXT9CT :5 9TRODUC$ GOATS DENUD$ ! >EA ( CACTUS PLANTS4 ,AN A4I;N 3C]N = SPECIALIZ$ ISL& ANIMALS IS T1 2C !Y H EVOLV$ 9 AN ECOSY/EM DEVOID ( -PETITORS & PR$ATORS1 !Y (T5 LOSE ! DEF5SIVE ME*ANISMS T O!R SPECIES TAKE = GRANT$4 ,= 9/.E1 ! 9TRODUC;N ( RATS & GOATS ONTO ! ,GALAPAGOS ,ISL&S HAS PROV$ DEVA/AT+ = NATIVE TORTOISE & L& IGUANA POPUL,NS4 ,! GOATS \-COMPETE ! TORTOISES = PLANT FOODS1 &! RATS FE$ ON _! E7S1 PREV5T+ !M F REPRODUC+4 ,= ^! SPECIES #CG 6BE A# 6SURVIVE 9 ! AGE ( MOD]N TRANSPORT,N1 HUMAN COLONIZ,N1 & JET AIRCRAFT1 !Y H 6BE VIGOR\SLY PROTECT$ 0MAN4 ,)\T A SU3ESS;L CAPTIVE BRE$+ PROGRAM &! /RICT REGUL,NS AB :O & :AT IS ALL[$ 6L& ON ! ,GALAPAGOS ,ISL&S1 ! NATIVE TORTOISES & L& IGUANAS WD SURELY G ! WAY (! DODO4 ,>TIFICIAL SELEC;N &! ,AGE (! ,E>? ,:5 ,D>W9 GOT BACK 6,5GL&1 HE SETTL$ D[N 6A QUIET LIFE 9 ! C.TRYSIDE & ASSEM#D 8 ?"\S4 ,Y'D ?9K HE WD H BE5 R1DY 6TELL ! _W AB 8 !ORIES1 B HE HELD BACK4 ,9 ,D>W9'S "D1 ! NO;N T EA* SPECIES 0 N SPECI,Y CR1T$ 0,GOD 0 HELD Z H]ETICAL1 & 9 PROMOT+ X1 HE WD H ISOLAT$ HMF F 8 COLL1GUES1 SUPPORT]S1 & SCI5TIFIC H]OES4 ,HE KNEW IF HE 0 GO+ 6PUBLI% S* AN #CH \TRAGE\S !ORY T HE WD NE$ 6H OV]:ELM+ EVID;E4 ,D>W9 SP5T ! NEXT TW5TY YE>S 3DUCT+ EXP]I;TS ON SE$S & B>NACLES -- HE WANT$ 6SEE IF PLANTS CD REPRODUCE AF _! SE$S _H BE5 AT SEA = MON?S1 & HE WANT$ 6DOCU;T ! HI< LEVEL ( V>I,N NEC = NATURAL SELEC;N 6O3UR 9 ANO!R TYPE ( ANIMAL4 ,D>W9 AL M>%AL$ EVID;E F DOME/IC SPECIES4 ,MEET+ ) E35TRIC PIGEON BRE$]S 9 DU/Y TAV]NS1 ,D>W9 >GU$ T IF PUNY HUMAN M$DL+ CD BR+ 96EXI/;E ! GRT V>IETY ( DOME/IC DOG & BIRD V>IETIES1 SURELY EONS ( NATURAL SELEC;N CD CR1TE ALL (! NATUR,Y O3URR+ V>IETIES ( PLANTS & ANIMALS4 ,? NO;N ( EONS BR+S UP AN IMPORTANT PO9T -- 9 ,D>W9'S "D X WASN'T WELL-E/ABLI%$ T ! _W 0 ALL T OLD4 ,A LIT]AL 9T]PRET,N (! BI# PUT ! #CI E>? AT ONLY #F1JJJ YE>S OLD1 N EV5 A FRAC;N (! "T NEC = BIOLOGICAL DIV]S;Y 6BE EXPLA9$ 0EVOLU;N Z ,D>W9 3CVD X4 ,M SCI5TIFIC,Y1 ! EM95T GEOLOGI/ ,WILLIAM ,?OMPSON1 LAT] "KN Z ,"L ,KELV91 0CALCULAT+ ! RATE ( COOL+ (! E>? F LAVA1 E/IMAT$ ! AGE (! E>? CD 2 NO GRT] ?AN #DJJ MILLION YE>S OLD4 ,? E/IMATE REMA9$ A MAJOR OBJEC;N 6A NATURAL EXPLAN,N ( BIOLOGICAL DIV]S;Y UNTIL ! 4COV]Y ( NUCLE> PHYSICS -- ! PRODUC;N ( H1T F ! BR1KD[N ( ELE;TAL "PICLES 9 ! E>?'S CORE ALL[S =A M* SL[] RATE ( COOL+1 &! ACTUATE #D.E BILLION YE> AGE (! E>?4 ,? HELPS 6EXPLA9 ! LACKLU/] SUPPORT = NATURAL SELEC;N 9 ! PA/ -- AL? ! 3CEPT ( EVOLU;N 9 G5]AL 0 A3EPT$ 0SCI5TI/S ALM IMMLY AF ! PUBLI%+ ( ,D>W9'S "W1 BIOLOGI/S ?"\ HE'D GOTT5 #DJ ! ME*ANISM WR;G WELL 96! #AICJ;S4 ,WALLACE & ,BIOGEOGRAPHY ,:ILE ,D>W9 0 PUTT]+ >.D 8 G>D5S M1SUR+ ! L5G? ( B>NACLE'S V>I\S ANATOMICAL NOVELTIES1 LL DID HE "K "! 0 ANO!R BRILLIANT AMATEUR NATURALI/ HOT ON ! TRAIL ( NATURAL SELEC;N4 ,ALFR$ ,RUSSELL ,WALLACE1 =C$ 6SUPPORT HMF )\T ! FAMILY 9H]IT.E T ,D>W9 5JOY$1 _H L$ AN EXCIT+ AN ADV5TUR\S LIFE FULL (! H>D%IP ( TROPICAL TRAVEL 9 ! ,VICTORIAN AGE4 ,3/ANTLY VICTIM 6RA%ES1 9SECT /+S1 & FREQU5T B\TS ( MAL>IA1 ,WALLACE N"O!.S COLLECT$ _M R>E & VALUA# SPECIM5S 9 ! ,AMAZON4 ,RETURN+ 6,5GL&1 8 %IP CAU4 ,J ACR A N>R[ /RAIBOR1L R1LM 0 FILL$ 9/1D 0! RELATIVELY QUIET MUN*+ ( TREE KANG>OOS4 ,WALLACE 0 AN 9TELLIG5T MAN1 9T5T ON G5]AT+ AN EXPLAN,N =! 8MY/]Y ( MY/]IES0 -- ! S\RCE ( NEW SPECIES ( LIFE ON ,E>?4 ,"W+ BACKW>DS1 ,WALLACE R1LIZ$ T ! ANIMALS EA/ (! MY/]I\S B.D>Y M/ 2 H ORIG9AT$ )! ,AU/RALIAN CR1TURES1 &! O!RS 9 ,ASIA4 ,! ISL&S M/ H BE5 #DB 3NECT$ 9 ! PA/1 )! OC1N WAT]S ONLY REC5TLY 9T]V5+4 ,! SPLIT AL;G ! L9E M/ 2 M* M SIGNIFICANT -- ^! ANIMALS _H 6H ORIG9AT$ 9DEP5D5TLY4 ,MOD]N GEOLOGY HAS BORNE \ ,WALLACE'S ?9K+1 REV1L+ T ! WAT] SEP>AT+ ! ISL&S ON EI SIDE ( 8 L9E IS M* DEEP] ?AN ! WAT]S 9 2T ! O!R ISL&S4 ,:5 WAT] LEVELS 7 L[] 9 ! PA/1 ! >EA 3SI/$ ( TWO L&MASSES 9/1D (A BUN* ( ISL&S4 ,WALLACE KNEW T ! 4TRIBU;N ( ANIMALS & PLANTS ON ^! ISL&S 9DICAT$ T ORGANISMS 7 RELAT$ 6EA* O!R1 & T !Y "!=E M/ H A -MON ANCE/OR1 B UNTIL AN ESPECI,Y VIRUL5T B\T ( MAL>IA 3F9$ HM 6HIS B$ ) NO?+ B 8 ?"\S1 HE CDN'T FIGURE \ ! ME*ANISM ( NATURAL SELEC;N4 ,AF 8 EPIPHANY1 HE S5T A DRAFT ( 8 ID1S 6,D>W9 : M/ H S5T ! ELD] NATURALI/ 96HY/]ICS -- 8 CAU;N MIW9'S 8BIG BOOK0 ..,ON ! ,ORIG9 ( .,SPECIES1 WD EVOLU;N 2COME A H\SEHOLD 3CEPT &! T>GET ( WIDESPR1D CRITICISM4 ,ISL& BIOGEOGRAPHY & 3S]V,N PRACTICE ,LET'S G BACK 6,C5TRAL ,P>K4 ,REMEMB]1 A P>K IS ESS5TI,Y AN ISL& ( WILD;S 9 ! OC1N ( URBAN DEVELOP;T4 ,US+ ! REL,N%IPS SCI5TI/S H 4COV]$ F R1L ISL&S1 WE C PREDICT :AT W HAPP5 6,C5TRAL ,P>K4 ,IF ! URBAN "P ( ,NEW ,YORK ,C;Y EXP&$1 ! IMMIGR,N RATE 6! P>K WD DECR1SE4 ,Z AN ISL&1 C5TRAL P>K IS PRETTY SMALL1 AB #HEJ ACRES1 R\K ISN'T TOT,Y WILD4 ,OV] ! 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